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This is a bug in the tree rule, since defining the tree_exclude_sites parameter should add --exclude-sites config/{lineage}/{segment}/exclude-sites.txt to the augur tree command. We only have those "exclude sites" files for HA and only the H1N1pdm file has anything in it. The loss of that functionality must be related to the refactored workflow.
Possible solution
We should update the tree rule in the core workflow rules to check for the tree_exclude_sites config and create appropriate augur tree argument for that file if it is provided.