_Yarrowia lipolytica_ should have complex I for respiratory chain, not the alternative internal NADH:ubiquinone oxidoreductase(NDI1 in _S. cervesiae_ ) `Reaction : y000773 NADH:ubiquinone oxidoreductase H+[m] + NADH[m] + ubiquinone-6[m] => NAD[m] + ubiquinol-6[m] 1.6.5.9 YALI0F25135g` should pump proton.