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A non-zero number of seeds germinated after 2 years instead of just 1 in Bruna 2002 (Fig 4). E.g., it looks like in continuous forest, and additional ~4% of seeds germinated in the second year. One way to model this could be to add another discrete class, seed
that has age related transitions into sdlg
.
Bruna, Emilio M. “Effects of Forest Fragmentation on Heliconia Acuminata Seedling Recruitment in Central Amazonia.” Oecologia 132, no. 2 (2002): 235–43. https://doi.org/10.1007/s00442-002-0956-y.
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